I know it doesn't look like classic iron def, which would usually start at the base of the leaf on new growth, so I'm probably way off here.
On the other hand, the more I see the pictures and go over everything, it really makes me think this may be a molybdenum deficiency. Probably cause by ph lockout. I think this could be one of the main issues, especially because its so hard to diagnose, as it looks kinda like N, Ca and Mg def.
Really hope this helps, not sure if anyone already said this, if so I apologize.